Cockatoo

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A cockatoo is any of the 21 bird species belonging to the family Cacatuidae. Along with the Psittacidae family (the true parrots), they make up the order Psittaciformes. The name cockatoo originated from the Malay name for these birds, kaka(k)tua (either from kaka "parrot" + tuwah, or "older sister" from kakak "sister" + tua, "old"^{[verification needed]}).

Cockatoos share many features with other parrots including the characteristic curved beak shape and a zygodactyl foot, with two forward toes and two backwards toes. They differ, however in a number of characteristics, including the often spectacular movable headcrest, the presence of a gall bladder and some other anatomical details, and their lack of the Dyck texture feather composition which causes the bright blues and greens seen in true parrots. Also Cockatoo species are, on average, larger than the average size of true parrots; however, the cockatiel, the smallest Cockatoo, is a small bird, and some of the largest parrots including the Hyacinth Macaw (the longest parrot) and the flightless Kakapo (the heaviest parrot), are true parrots. Placement of the cockatoos as a separate family is fairly undisputed, but it is not resolved whether or not other living lineages of parrots (such as the lories and lorikeets) are as distinct as they are.

Cockatoos have a much more restricted range than the true parrots, occurring naturally only in Australia and nearby islands. Eleven of the 21 species exist in the wild only in Australia, while seven species occur in Indonesia, New Guinea, and other south Pacific islands. Three species occur in both New Guinea and Australia.

[edit] Cockatoos as endangered or vulnerable species

All species of cockatoo are protected by the Convention on International Trade in Endangered Species of Wild Fauna and Flora (also known as CITES), which makes the import, export and trade in all wild-caught parrots and cockatoos illegal.

The following cockatoo species are protected on the CITES appendix 1 list of endangered species.

• Goffin's cockatoo, Cacatua goffini
• Red-vented Cockatoo, Cacatua haematuropygia
• Moluccan Cockatoo, Cacatua moluccensis
• Yellow-crested Cockatoo, Cacatua sulphurea
  • includes the subspecies, Citron-crested Cockatoo, Cacatua sulphurea citrinocristata
• Palm Cockatoo, Probosciger aterrimus

All of the other cockatoo species are protected on the CITES appendix 2 list of vulnerable species.

[edit] Systematics and evolution

Brown & Toft (1999) reviewed the existing evidence and additional mitochondrial 12S rRNA sequence data to arrive at a well-supported phylogeny of the cockatoos. They could distinguish 3 subfamilies:

1. The all-black Palm Cockatoo represents distinct lineage that diverged early; it was previously sometimes grouped with the other black species but this is incorrect.
2. The dark cockatoos; sexually dichromatic species which have ample melanin in their plumage and some red, yellow or orange on wing, tail and face, barred feathers on wing, tail and/or body as well as contrasting ear area spotting in females, while males have the corresponding feathers unbarred and may lack the ear spotting. This group includes the remaining black cockatoos, the Gang-gang Cockatoo and, interestingly, the cockatiel which had previously been placed in a subfamily of its own (Nymphicinae) or even as a broad-tailed parrot.
3. The remaining species, which are all hypomelanistic and not sexually dimorphic.

The genera Calyptorhynchus and Cacatua can be further resolved into two subgenera each, and in the latter case as a distinct third lineage the white-and-pink Major Mitchell's Cockatoo, which is intermediate in coloration between the grey-and-pink Galah and the white Cacatua. It is best recognized as a monotypic genus Lophocroa. Indeed, pending further research, all subgenera could conceivably be raised to genus rank.

Intron 7 of nuclear β-fibrinogen sequence data suggests that the Microglossinae may have diverged later, and that the cockatiel might be distinct enough to warrant recognition of the Nymphicinae (Astuti, 2004?), but in other aspects agrees with the rRNA and newly-interpreted morphological data.

The fossil record of cockatoos is even more limited than that of parrots in general, with only one truly ancient cockatoo fossil known: A species of Cacatua, most probably subgenus Licmetis, was found in Early Miocene (16-23 mya) deposits of Riversleigh, Australia (Boles, 1993). In Melanesia, subfossil bones of Cacatua species which apparently did not survive early human settlement were found on New Caledonia and New Ireland^{[citation needed]}. The bearing of these fossils on cockatoo evolution and phylogeny is fairly limited, except that the Riversleigh fossil allows some tentative dating of the divergence of subfamilies.

[edit] Family Cacatuidae

• Subfamily Microglossinae
  • Genus Probosciger
    • Palm Cockatoo, Probosciger aterrimus
• Subfamily Calyptorhynchinae - dark cockatoos
  • Genus Callocephalon
    • Gang-gang Cockatoo, Callocephalon fimbriatum
  • Genus Nymphicus (tentatively placed here)
    • Cockatiel, Nymphicus hollandicus
  • Genus Calyptorhynchus
    • Subgenus Calyptorhynchus - black-and-red cockatoos
      • Red-tailed Black Cockatoo, Calyptorhynchus (Calyptorhynchus) banksii
      • Glossy Black Cockatoo, Calyptorhynchus (Calyptorhynchus) lathami
    • Subgenus Zanda - black-and-yellow/white cockatoos
      • Yellow-tailed Black Cockatoo, Calyptorhynchus (Zanda) funereus
      • Short-billed Black Cockatoo, Calyptorhynchus (Zanda) latirostris
      • Long-billed Black Cockatoo, Calyptorhynchus (Zanda) baudinii
  • Genus Eolophus
    • Galah, Eolophus roseicapilla
• Subfamily Cacatuinae - white cockatoos
  • Genus Cacatua
    • Subgenus Cacatua - true white cockatoos
      • Major Mitchell's Cockatoo (also called Leadbeater's Cockatoo), Cacatua leadbeateri
      • Sulphur-crested Cockatoo, Cacatua (Cacatua) galerita
      • Yellow-crested Cockatoo, Cacatua (Cacatua) sulphurea
        • Citron-crested Cockatoo, Cacatua (Cacatua) sulphurea citrinocristata
      • Blue-eyed Cockatoo, Cacatua (Cacatua) ophthalmica
      • Moluccan Cockatoo (also called Salmon-crested Cockatoo), Cacatua (Cacatua) moluccensis
      • Umbrella Cockatoo, Cacatua (Cacatua) alba
    • Subgenus Licmetis - corellas
      • Long-billed Corella, Cacatua (Licmetis) tenuirostris
      • Western Corella, Cacatua (Licmetis) pastinator
      • Little Corella, Cacatua (Licmetis) sanguinea
      • Red-vented Cockatoo, Cacatua (Licmetis) haematuropygia
      • Goffin's Cockatoo, Cacatua (Licmetis) goffini
      • Ducorps' Cockatoo, Cacatua (Licmetis) ducorpsii

[edit] Popular culture

Cockatoos often have pronounced responses to musical sounds, and numerous videos exist showing the birds "dancing" to popular music, although it is not known if their musical perception is any different from that of other birds. A search with the terms "dancing cockatoo" yields over 200 results on YouTube, mostly by private owners.

[edit] References

• Astuti, Dwi (2004?): A phylogeny of cockatoos (Aves: Psittaciformes) inferred from DNA sequences of the seventh intron of nuclear β-fibrinogen gene. Doctoral work, Graduate School of Environmental Earth Science, Hokkaido University, Japan. PDF fulltext
• Boles, W.E. (1993): A new cockatoo (Psittaciformes: Cacatuidae) from the Tertiary of Riversleigh, northwestern Queensland, and an evaluation of rostral characters in the systematics of parrots. Ibis 135: 8-18.
• Brown, D.M. & Toft, C.A. (1999): Molecular systematics and biogeography of the cockatoos (Psittaciformes: Cacatuidae). Auk 116(1): 141-157.

[edit] External links

• MyToos.com - site warning against cockatoo ownershipbg:Какаду

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